1. INRA, Laboratoire des Interactions Plantes–Microorganismes (LIPM), UMR441, Chemin de Borde Rouge F‐31326 Castanet‐Tolosan, France
2. CNRS, Laboratoire des Interactions Plantes–Microorganismes (LIPM), UMR2594, Chemin de Borde Rouge F‐31326 Castanet‐Tolosan, France
3. Department of International Cooperation, Chinese Academy of Agricultural Sciences, 12 Zhongguancun South Street, Beijing 100081, China
4. College of Biological Sciences, China Agricultural University, Beijing 100193, China