1. Department of Genetics, Faculty of Science, Kasetsart University, 50 Ngam Wong Wan Road, Chatuchak, Bangkok 10900, Thailand
2. Center for Advanced Studies in Tropical Natural Resources, Kasetsart University, Ngam Wong Wan, Chatuchak, Bangkok 10900, Thailand
3. CNAP, Biology Department, University of York, Heslington, York YO10 5DD, UK