1. Laboratory of Genetics, Wageningen University, Droevendaalsesteeg 1, 6708 PB Wageningen, the Netherlands
2. Biosystematics Group, Wageningen University, Droevendaalsesteeg 1, 6708 PB Wageningen, the Netherlands
3. Wageningen UR Plant Breeding, Droevendaalsesteeg 1, 6708 PB Wageningen, the Netherlands