1. Netherlands Institute of Ecology (NIOO‐KNAW), PO Box 50, 6708 PB Wageningen, the Netherlands
2. Centre for Estuarine and Marine Ecology, Netherlands Institute of Ecology (NIOO‐KNAW), PO Box 140, 4400 AC Yerseke, the Netherlands
3. Insititute of Biology, Leiden University, Leiden, the Netherlands