1. NERC Unit of Comparative Plant Ecology, Department of Animal and Plant Sciences, The University, Sheffield S10 2TN, UK
2. Horticulture Research International, Wellesbourne, Warwick CV35 9EF, UK
3. Department of Chemistry, The University, Sheffield S10 2TN, UK
4. Department of Environmental Biology, University of Manchester, Manchester M13 9PL, UK