1. National Institute of Livestock and Grassland Science, Nishi‐nasuno, Tochigi, 329‐2793 Japan;
2. Present address: The University of Tokyo, Biotechnology Research Center, 1‐1‐1 Yayoi, Bunkyo‐ku, Tokyo 113‐8657, Japan;
3. The Samuel Roberts Noble Foundation, Plant Biology Division, 2510 Sam Noble Parkway, Ardmore, Oklahoma 74301, USA;
4. Present address: Boyce Thompson Institute, Tower Road, Ithaca, New York 14853, USA;
5. Present address: National Institute for Agro‐Environmental Sciences, Tsukuba, Ibaraki 305‐8604, Japan