1. Institute of Grassland and Environmental Research, Plas Gogerddan, Aberystwyth, Ceredigion SY23 3EB, UK;
2. Institute of Biological Sciences, Aberystwyth University, Ceredigion SY23 3DA, UK;
3. Instituto de la Grasa (CSIC), Avda Padre García Tejero 4, Sevilla 41012, Spain;
4. Department of Computer Science, Aberystwyth University, Ceredigion SY23 3DB, UK