1. Soil Science Department, Rothamsted Research, Harpenden, Hertfordshire AL5 2JQ, UK;
2. Research Institute for Bioresources, Okayama University, Chuo 2‐20‐1, Kurashiki 710‐0046, Japan;
3. School of Biological Sciences, University of Aberdeen, Cruickshank Building, St Machar Drive, Aberdeen AB24 3UU, UK