1. Weed Science Center, Utsunomiya University, 350 Mine‐machi, Utsunomiya 321‐8505, Japan;
2. Faculty of Agriculture, Utsunomiya University, 350 Mine‐machi, Utsunomiya 321‐8505, Japan;
3. Graduate School of Life and Environmental Sciences, Osaka Prefecture University, 1‐1 Gakuencho, Naka‐ku, Sakai, Osaka 599‐8531, Japan