1. Earth and Environmental Sciences Division, Los Alamos National Laboratory, Los Alamos, NM 87545, USA;
2. Department of Biology, MSC03 2020, 1 University of New Mexico, Albuquerque, NM 87131‐0001, USA;
3. US Geologcial Survey, Jemez Mountains Field Station, 15 Entrance Road, Los Alamos, NM 87544, USA;
4. School of Natural Resources, Institute for the Study of Planet Earth, and Department of Ecology and Evolutionary Biology, University of Arizona, Tucson, AZ 85721‐0043, USA;
5. Merriam‐Powell Center for Environmental Research, Peterson Hall, Bldg 22, Rm 330, Box 6077, Northern Arizona University Flagstaff, AZ 86011, USA;
6. School of Forestry, Northern Arizona University, Flagstaff, AZ 86001‐5018, USA;
7. Department of Biology, University of Utah, 257S 1400E, Salt Lake City, UT 84112, USA;
8. Department of Integrative Biology, University of California, Berkeley, CA 94720;
9. Botany Department, University of Cape Town, Private Bag, Rondebosch, 7700, South Africa;
10. Department of Renewable Resources, University of Wyoming, Laramie, WY 82071 USA;
11. Department of Biology, University of New Mexico, Albuquerque, NM 87131, USA