1. Centre de Recherche Nestlé, 101 Av. Gustave Eiffel, Notre Dame d’Oé, BP 49716‐37097 Tours, France;
2. School of Agriculture, Food and Rural Development, Newcastle University, Newcastle upon Tyne NE1 7RU, UK
3. Department of Plant Breeding, 248 Emerson, Cornell University, Ithaca, NY 14853, USA;