1. Department of Plant and Microbial Biology and
2. Environmental Science Policy and Management; University of California, Berkeley, CA 94720‐3102, USA;
3. Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA, 02138, USA;
4. USDA‐ARS, Department of Plant Sciences, University of Arizona, PO Box 210036, Tucson, AZ 85721, USA