1. Plant, Soil, and Insect Sciences, University of Massachusetts, Amherst, MA 01003, USA;
2. Present address: Entomology, Soils & Plant Sciences, Clemson University, Clemson SC 29634, USA
3. Biology Department, University of Massachusetts, Amherst, MA 01003, USA;
4. School of Natural Science, Hampshire College, Amherst, MA 01002, USA;