1. Biology and Physical Geography Unit and SARAHS Centre, University of British Columbia Okanagan, Kelowna, BC V1V 1V7, Canada
2. Department of Forest Sciences, University of British Columbia, Vancouver, BC V6T 1Z4, Canada
3. Department of Biology, University of Western Ontario, London, ON N6A 5B8, Canada
4. SUNY College of Environmental Science and Forestry, Faculty of Environmental and Forest Biology, One Forestry Drive, Syracuse, NY 13210‐2788, USA