1. Forest Ecology and Forest Management Group, Centre for Ecosystem Studies, Wageningen University, PO Box 47, 6700 AA Wageningen, the Netherlands
2. Instituto Boliviano de Investigación Forestal (IBIF), Casilla 6204, Santa Cruz, Bolivia
3. Resource Ecology Group, Centre for Ecosystem Studies, Wageningen University, PO Box 47, 6700 AA Wageningen, the Netherlands
4. Department of Crop Sciences, Agronomy in the Tropics, Georg‐August‐Universität Göttingen, Grisebachstrasse 6, 37077 Göttingen, Germany