1. Laboratory of Plant Systematics, K.U. Leuven, Kasteelpark Arenberg 31, B‐3001 Leuven, Belgium
2. Department of Plant & Microbial Biology, University of California Berkeley, Berkeley, CA 94720, USA
3. Department of Evolution, Ecology, and Organismal Biology, The Ohio State University Herbarium, 1315 Kinnear Road, Columbus, OH 43212, USA