1. Department of Plant and Microbial Biology, University of California, Berkeley, CA 94720, USA
2. Department of Biology, Lewis and Clark College, Portland, OR 97219, USA
3. Center for Tropical Forest Science, Smithsonian Tropical Research Institute and Arnold Arboretum, Harvard University, Cambridge, MA 02138, USA
4. Forest Research Centre, Sarawak Forestry Corporation, Kuching, Sarawak, Malaysia & Center for Tropical Forest Science‐Arnold Arboretum Asia Program, Cambridge, MA 02138, USA