1. ECORES lab, Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, China
2. Department of Botany, College of Horticulture and Landscape Architecture, Southwest University, Chongqing 400716, China
3. Department of Plant Biology, Cornell University, Ithaca, NY 14850, USA
4. Department of Biology, Nanjing University, 22 Hankou Road, Nanjing 210093, China