1. Theoretical Research in Evolutionary Life Sciences, Groningen Institute for Evolutionary Life SciencesUniversity of Groningen 9747 AG Groningen The Netherlands
2. Laboratorio de Biología Evolutiva de Vertebrados, Departamento de Ciencias BiológicasUniversidad de los Andes Bogotá Colombia
3. Department of Zoology and Biodiversity Research CentreUniversity of British Columbia Vancouver British Columbia V6T 1Z4 Canada