1. Fisheries Department, Faculty of Natural Resources University of Guilan Sowmeh Sara Iran
2. Department of Marine Sciences, The Caspian Sea Basin Research Center University of Guilan Rasht Iran
3. Department of Fisheries and Environmental Sciences Gorgan University of Agricultural Sciences and Natural Resources Gorgan Iran
4. Institute of Life, Earth and Environment (ILEE), Research Unit in Environmental and Evolutionary Biology University of Namur Namur Belgium