1. Zoological Institute; University of Basel; Vesalgasse 1 Basel CH-4051 Switzerland
2. Centre d'Ecologie Fonctionnelle et Evolutive - UMR 5175; campus CNRS 1919 route de Mende 34293 Montpellier Cedex 5 France
3. Department of Biology; Ecology and Evolution; University of Fribourg; Chemin du Muśee 10 1700 Fribourg Switzerland