1. Marine Spatial Ecology Laboratory and Australian Research Council Centre of Excellence for Coral Reef Studies, School of Biological Sciences, Goddard Building, The Univ. of Queensland Brisbane QLD 4072 Australia
2. Southern Seas Ecology Laboratories, School of Biological Sciences and The Environment Institute DX 650 418, The Univ. of Adelaide Adelaide SA 5005 Australia
3. Dept of Animal Ecology and Ecophysiology Inst. for Water and Wetland Research, Radboud University Nijmegen Toernooiveld 1 NL‐6525 ED Nijmegen the Netherlands
4. College of Life and Environmental Sciences, Univ. of Exeter Stocker Road Exeter EX4 4QD UK