1. Kawatabi Field Science Centre, Graduated School of Agricultural Science Tohoku University Osaki Miyagi Japan
2. Graduate School of Agriculture Kyoto University Kyoto Japan
3. Systematic and Evolutionary Botany and Biodiversity Group, MOE Laboratory of Biosystem Homeostasis and Protection, College of Life Sciences, Zhejiang University Hangzhou 310058 China
4. Graduate School of Human and Environmental Studies Kyoto University Kyoto Japan
5. Research Institute of Energy, Environment and Geology, Hokkaido Research Organization Sapporo Hokkaido Japan