1. Department of Bioscience, Graduate School of Agriculture Ehime University 3‐5‐7 Tarumi Matsuyama Ehime 790‐8566Japan
2. Faculty of Agriculture Ehime University 3‐5‐7 Tarumi Matsuyama Ehime 790‐8566Japan
3. Center for Marine Environmental Studies Ehime University 2‐5 Bunkyo‐cho Matsuyama Ehime 790‐8577Japan
4. Faculty of Food and Nutritional Sciences Toyo University 1‐1‐1 Izumino, Itakura‐machi Ora‐gun Gunma 374‐0193Japan