1. Tianjin Key Laboratory of Animal and Plant Resistance College of Life Sciences, Tianjin Normal University Tianjin 300387 China
2. State Key Laboratory of Plant Genomics and National Center for Plant Gene Research Institute of Genetics and Developmental Biology, the Chinese Academy of Sciences Beijing 100101 China
3. CAS Center for Excellence in Molecular Plant Sciences, the Chinese Academy of Sciences Beijing 100101 China
4. University of Chinese Academy of Sciences Beijing 100049 China