1. Department of Biology and Botanic Garden University of Fribourg Fribourg Switzerland
2. Department of Geosciences University of Fribourg Fribourg Switzerland
3. Info Flora, the National Data and Information Center on the Swiss Flora Bern Switzerland
4. Swiss Federal Institute of Forest, Snow and Landscape Research WSL Birmensdorf Switzerland
5. Eastern China Conservation Centre for Wild Endangered Plant Resources, Shanghai Chenshan Botanical Garden Shanghai China
6. Natural History Museum Fribourg Fribourg Switzerland