1. Department of Natural History Sciences, Graduate School of Science Hokkaido University North 10, West 8 060‐0810 Sapporo Japan
2. Faculty of Environmental Earth Science Hokkaido University North 10 West 5 060‐0810 Sapporo Japan
3. Department of Biological Sciences, Faculty of Science Hokkaido University North 10, West 8 060‐0810 Sapporo Japan