1. Australian Centre for Ancient DNA, School of Earth and Environmental Sciences; University of Adelaide; Adelaide; SA; 5005; Australia
2. Department of Biology; University of Oslo; PO Box 1066; Blindern, NO-0316; Oslo; Norway
3. Department of Earth Science; University of Bergen; Allegaten 41, NO-5007; Bergen; Norway
4. Centre for GeoGenetics, Natural History Museum of Copenhagen; University of Copenhagen; Universitetsparken 15, DK-2100; Copenhagen Ø; Denmark
5. Centro Mixto; Universidad Complutense de Madrid-Instituto de Salud Carlos III de Evolucion y Comportamiento Humanos; c/Sinesio Delgado 4 Pabellon 14; 28029; Madrid; Spain