1. Key Laboratory of Tropical Forest Ecology Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Kunming Yunnan Province 650223 China
2. College of Bioscience and Biotechnology Shenyang Agricultural University Shenyang Liaoning Province 110866 China
3. Division of Biological Sciences and the Institute on Ecosystems The University of Montana Missoula MT 59812 USA
4. Departamento de Ecologôa de la Biodiversidad Instituto de Ecología Universidad Nacional Autônoma de Mexico Apartado Postal 70‐275 CP 04510 México DF Mexico
5. College of Life Sciences Hebei University Baoding Hebei Province 071002 China
6. Department of Life Sciences University of the West Indies St Augustine Trinidad and Tobago