1. U.S. Geological Survey, Utah Cooperative Fish and Wildlife Research Unit, Department of Watershed Sciences & The Ecology Center; Utah State University; Logan Utah 84322-5210 U.S.A.
2. Department of Wildland Resources; Utah State University; Logan UT 84322-5230 U.S.A.
3. Department of Watershed Sciences; Utah State University; Logan Utah 84322-5210 U.S.A.