1. Imperial College London, Division of Biology; Silwood Park Campus, Manor House; Buckhurst Road, Ascot Berks SL5 7PY United Kingdom
2. Institute of Zoology; Zoological Society of London; Regents Park London NW1 4RY United Kingdom
3. University of Ghana; Legon Accra Ghana
4. Ghana Wildlife Division; Forestry Commission; Accra Ghana