1. Department of Biology University of Oxford Oxford OX1 3RB UK
2. State Key Laboratory of Plant Cell and Chromosome Engineering, Institute of Genetics and Developmental Biology Chinese Academy of Sciences Beijing 100101 China
3. National Key Laboratory of Crop Genetics & Germplasm Enhancement and Utilization Nanjing Agricultural University Nanjing 210095 China
4. College of Life Sciences University of Chinese Academy of Sciences Beijing 100049 China
5. New Cornerstone Science Laboratory Beijing 100101 China