1. Research Center for Environmental Bioscience; Graduate School of Science and Technology; Kwansei-Gakuin University; Sanda Hyogo 669-1337 Japan
2. Department of Bioscience; Graduate School of Science and Technology; Kwansei-Gakuin University; 2-1 Gakuen Sanda Hyogo 669-1337 Japan
3. Department of Biotechnology; College of Life Sciences; Ritsumeikan University; Kusatsu Shiga 525-8577 Japan