1. Centre d'Ecologie Fonctionnelle et Evolutive UMR 5175 CNRS – Université de Montpellier – Université Paul Valéry Montpellier – EPHE 1919 Route de Mende 34293 Montpellier Cedex 5 France
2. INRA UR 874 UREP, Grassland Ecosystem Research 5 Chemin de Beaulieu 63039 Clermont‐Ferrand France
3. INRA UMR AMAP Boulevard de la Lironde 34398 Montpellier Cedex 5 France
4. Institute of Biological Sciences University of Malaya 50603 Kuala Lumpur Malaysia
5. State Key Laboratory for Conservation and Utilization of Subtropical Agro‐bioresources, and College of Forestry Guangxi University Nanning Guangxi 530004 China