1. School of Clinical Medicine, Cambridge Institute for Medical Research, University of Cambridge, Cambridge, CB2 0XY, UK
2. Current address: Centre for Molecular Microbiology and Infection, Division of Investigative Sciences, Imperial College London, London, SW7 2AZ, UK
3. Department of Human Genetics, Center for Neurodegenerative Diseases, Emory University School of Medicine, Atlanta, GA 30322, USA