1. University of California, Irvine, Department of Earth System Science, Irvine, CA 92697-3100, U.S.A.;
2. Woods Hole Oceanograhic Institution, Department of Marine Chemistry and Geochemistry, MS #25, Woods Hole, MA 02543-1543, U.S.A.;
3. National Center for Atmospheric Research, P.O. Box 3000, Bouldery, CO 80307-3000, U.S.A.;
4. University of Southern California, Biology Department, Los Angeles, CA, 90089-0371, U.S.A.