1. Department of Aquaculture; National Taiwan Ocean University; No. 2, Peining Rd 20224 Keelung Taiwan
2. National Marine Fisheries Service National Systematics Laboratory; NOAA, Smithsonian Institution P. O. Box 37012 National Museum of Natural History WC-57, MRC-153; Washington DC 20013-7012 U.S.A.
3. Laboratory of Fish Ecology and Evolution; Biodiversity Research Center, Academia Sinica; Nankang 11529 Taipei Taiwan