1. CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany Chinese Academy of Sciences Kunming 650204 China
2. Florida Museum of Natural History University of Florida Gainesville Florida 32611‐7800 USA
3. Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Mengla 666303 Yunnan China
4. University of the Chinese Academy of Sciences Beijing 100049 China
5. Faculty of Land Resource Engineering Kunming University of Science and Technology Kunming 650093 China