1. College of Life and Environmental SciencesHangzhou Normal University Hangzhou 311121 China
2. Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of BotanyChinese Academy of Sciences Kunming 650201 China
3. Key Laboratory of Tropical Plant Resources and Sustainable Use, Xishuangbanna Tropical Botanical GardenChinese Academy of Sciences Mengla 666303 Yunnan China
4. Plant Germplasm and Genomics Center, Germplasm Bank of Wild Species, Kunming Institute of BotanyChinese Academy of Sciences Kunming 650201 China
5. Department of Life SciencesNatural History Museum London SW7 5BD UK
6. Center for Integrative Conservation, Xishuangbanna Tropical Botanical GardenChinese Academy of Sciences Mengla 666303 Yunnan China