1. State Key Laboratory of Grassland Agro‐Ecosystems College of Pastoral Agriculture Science and Technology Institute of Innovation Ecology Lanzhou University Lanzhou 730000 China
2. Institute of Ecology College of Urban and Environmental Sciences Key Laboratory for Earth Surface Processes of the Ministry of Education Peking University Beijing 100871 China
3. Key Laboratory of Adaptation and Evolution of Plateau Biota Northwest Institute of Plateau Biology Chinese Academy of Sciences Xining 810008 China
4. State Key Laboratory of Vegetation and Environmental Change Institute of Botany Chinese Academy of Sciences Beijing 100093 China
5. School of Biological Sciences Georgia Institute of Technology Atlanta Georgia 30332 USA
6. Environmental Program Rubenstein School of Environment and Natural Resources University of Vermont Burlington VT 05405 USA
7. Rubenstein School of Environment and Natural Resources University of Vermont Burlington VT 05405 USA
8. Gund Institute for Environment University of Vermont Burlington VT 05405 USA