1. INRA, UMR 1349, Institute of Genetics, Environment and Plant Protection; Domaine de la Motte; BP 35327, 35653; Le Rheu Cedex; France
2. CNRS - INRA, UMR CBGP (INRA - IRD - CIRAD - Montpellier SupAgro); Campus International de Baillarguet; CS 30016, F-19 34988; Montferrier sur Lez Cedex; France