1. Research Institute for Microbial Diseases; Osaka University; Suita Osaka 565-0871
2. Graduate School of Medicine; Osaka University; Suita Osaka 565-0871 Japan
3. School of Environmental and Life Science; University of Newcastle; Callaghan New South Wales 2308 Australia
4. School of Pharmaceutical Sciences; Osaka University; Suita Osaka 565-0871 Japan