1. Shanghai Universities Key Laboratory of Marine Animal Taxonomy and Evolution Shanghai Ocean University Shanghai 201306 China
2. Institute of Evolution and Marine Biodiversity Ocean University of China Qingdao 266003 China
3. Key Laboratory of Coastal Environmental Processes and Ecological Remediation Yantai Institute of Coastal Zone Research Chinese Academy of Science Yantai 264003 China
4. Shanghai Collaborative Innovation for Aquatic Animal Genetics and Breeding Shanghai Ocean University Shanghai 201306 China
5. National Demonstration Center for Experimental Fisheries Science Education Shanghai Ocean University Shanghai 201306 China