1. Borneo Marine Research Institute Universiti Malaysia Sabah Jalan UMS Kota Kinabalu, Sabah Malaysia
2. Higher Institution Center of Excellence (HICoE) Institute of Tropical Aquaculture and Fisheries Universiti Malaysia Terengganu Kuala Nerus Terengganu Malaysia
3. Department of Marine Biosciences Tokyo University of Marine Science and Technology Konan Minato, Tokyo 108‐8477 Japan
4. Maizuru Fisheries Research Station Field Science Education and Research Center Kyoto University Maizuru Kyoto 625‐0086 Japan