1. Conservation Ecology Group Groningen Institute for Evolutionary Life Sciences (GELIFES) University of Groningen P.O. Box 11103 9700 CC Groningen The Netherlands
2. Department of Biological Sciences University of South Carolina 715 Sumter Street Columbia SC 29208 USA
3. NIOZ Royal Netherlands Institute for Sea Research Department of Coastal Systems and Utrecht University P.O. Box 59 1790 AB Den Burg Texel The Netherlands