1. School of Biological Sciences; University of Essex; Wivenhoe Park; Colchester; CO4 3SQ; UK
2. Equipe Environnement et Microbiologie; UMR CNRS-IPREM 5254; IBEAS; Université de Pau et des Pays de l'Adour; Pau; BP 1155 F-64013; France
3. Department of Biological Oceanography; Royal Netherlands Institute for Sea Research (NIOZ); PO Box 50; NL 1790; AB Den Burg; The Netherlands