1. School of Biological Sciences; Royal Holloway University of London; Egham Surrey TW20 0EX UK
2. Royal Botanic Gardens; Kew, Richmond Surrey TW9 3AB UK
3. Forest Research; Alice Holt; Farnham Surrey GU10 4LH UK
4. INRA; UMR 1202 BIOGECO; F-33610 Cestas France
5. BIOGECO; UMR 1202; Univ. Bordeaux; F-33400 Talence France