1. Department of Biochemistry; Hamamatsu University School of Medicine; Hamamatsu Japan
2. Laboratory of Developmental Genetics, Center for Experimental Medicine and Systems Biology, Institute of Medical Science; The University of Tokyo; Tokyo Japan
3. Section of Molecular Craniofacial Embryology, Graduate School of Medical and Dental Sciences; Tokyo Medical and Dental University; Tokyo Japan
4. Erin Lambers, Feinberg Cardiovascular Research Institute, Department of Medicine; Northwestern University School of Medicine; Chicago Illinois USA
5. Stowers Institute for Medical Research; Kansas City Missouri USA
6. Department of Anatomy and Cell Biology; University of Kansas Medical Center; Kansas City Missouri USA