1. Department of Integrative Biology; University of California; Berkeley CA 94720 USA
2. Department of Biology; Faculty of Science; Shinshu University; 3-1-1 Asahi Matsumoto Nagano 390-8621 Japan
3. Graduate School of Life and Environmental Science; Osaka Prefecture University; 1-1 Gakuen-cho Nakaku Sakai Osaka 599-8531 Japan
4. Division of Evolution; Ecology & Genetics; Research School of Biology; The Australian National University; Acton Canberra ACT 2601 Australia
5. Corporación Colombiana de Investigación Agropecuaria (CORPOICA); Centro de Investigación Palmira; Calle 23, Carrera 37, Continuo al Penal, Palmira Valle Colombia
6. Graduate School of Fisheries and Environmental Sciences; Nagasaki University; 1-14 Bunkyo-machi Nagasaki 852-8521 Japan
7. Graduate School of Human and Environmental Studies; Kyoto University; Yoshida-honmachi, Sakyo- ku, Kyoto 606-8501 Japan
8. Graduate School of Natural Sciences, and Research Center for Biological Diversity; Nagoya City University; 1 Yamanohata, Mizuho-cho, Mizuho-ku, Nagoya Aichi 467-8501 Japan
9. Institute of Mountain Science; Shinshu University; 3-1-1 Asahi, Matsumoto Nagano 390-8621 Japan