1. Soil and Ecosystem Ecology Laboratory; Lancaster Environment Centre; Lancaster University; Lancaster LA1 4YQ UK
2. Landcare Research; Lincoln 7640 Canterbury New Zealand
3. Laboratoire d'Ecologie Microbienne; Université Lyon1; Université de Lyon; USC INRA 1364; UMR CNRS 5557; 69622 Villeurbanne Cedex France
4. Faculty of Life Sciences; The University of Manchester; Michael Smith Building Oxford Road Manchester M13 9PT UK